<?xml version="1.0" encoding="UTF-8"?>
<!DOCTYPE article PUBLIC "-//NLM//DTD JATS (Z39.96) Journal Publishing DTD v1.3 20210610//EN" "JATS-journalpublishing1-3.dtd">
<article article-type="research-article" dtd-version="1.3" xmlns:mml="http://www.w3.org/1998/Math/MathML" xmlns:xlink="http://www.w3.org/1999/xlink" xmlns:xsi="http://www.w3.org/2001/XMLSchema-instance" xml:lang="ru"><front><journal-meta><journal-id journal-id-type="publisher-id">aari</journal-id><journal-title-group><journal-title xml:lang="ru">Проблемы Арктики и Антарктики</journal-title><trans-title-group xml:lang="en"><trans-title>Arctic and Antarctic Research</trans-title></trans-title-group></journal-title-group><issn pub-type="ppub">0555-2648</issn><issn pub-type="epub">2618-6713</issn><publisher><publisher-name>Государственный научный центр Российской Федерации Арктический и антарктический научно-исследовательский институт</publisher-name></publisher></journal-meta><article-meta><article-id pub-id-type="doi">10.30758/0555-2648-2026-72-2-236-251</article-id><article-id custom-type="elpub" pub-id-type="custom">aari-858</article-id><article-categories><subj-group subj-group-type="heading"><subject>Research Article</subject></subj-group><subj-group subj-group-type="section-heading" xml:lang="ru"><subject>ЭКОЛОГИЯ, БИОЦЕНОЛОГИЯ И БИОГЕОГРАФИЯ</subject></subj-group><subj-group subj-group-type="section-heading" xml:lang="en"><subject>ECOLOGY, BIOCENOLOGY AND BIOGEOGRAPHY</subject></subj-group></article-categories><title-group><article-title>Особенности социоприродных взаимодействий и динамика влияния оленеводства на окружающую среду на севере Западной Сибири во второй половине XIX — первой четверти XXI в.</article-title><trans-title-group xml:lang="en"><trans-title>Features of socio-natural interactions and the dynamics of the environmental impact of reindeer herding in the north of Western Siberia in the second half of the 19th century and the first quarter of the 21st century</trans-title></trans-title-group></title-group><contrib-group><contrib contrib-type="author" corresp="yes"><contrib-id contrib-id-type="orcid">https://orcid.org/0000-0002-7771-4592</contrib-id><name-alternatives><name name-style="eastern" xml:lang="ru"><surname>Соромотин</surname><given-names>А. В.</given-names></name><name name-style="western" xml:lang="en"><surname>Soromotin</surname><given-names>A. V.</given-names></name></name-alternatives><bio xml:lang="ru"><p>Тюмень</p></bio><bio xml:lang="en"><p>Tyumen</p></bio><email xlink:type="simple">a.v.soromotin@utmn.ru</email><xref ref-type="aff" rid="aff-1"/></contrib><contrib contrib-type="author" corresp="yes"><contrib-id contrib-id-type="orcid">https://orcid.org/0000-0001-6338-7669</contrib-id><name-alternatives><name name-style="eastern" xml:lang="ru"><surname>Московченко</surname><given-names>Д В.</given-names></name><name name-style="western" xml:lang="en"><surname>Moskovchenko</surname><given-names>D. V.</given-names></name></name-alternatives><bio xml:lang="ru"><p>Тюмень</p></bio><bio xml:lang="en"><p>Tyumen</p></bio><xref ref-type="aff" rid="aff-1"/></contrib><contrib contrib-type="author" corresp="yes"><contrib-id contrib-id-type="orcid">https://orcid.org/0000-0002-3658-746X</contrib-id><name-alternatives><name name-style="eastern" xml:lang="ru"><surname>Федоров</surname><given-names>Р. Ю.</given-names></name><name name-style="western" xml:lang="en"><surname>Fedorov</surname><given-names>R. Yu.</given-names></name></name-alternatives><bio xml:lang="ru"><p>Тюмень</p><p>Москва</p></bio><bio xml:lang="en"><p>Tyumen</p><p>Moscow</p></bio><xref ref-type="aff" rid="aff-2"/></contrib><contrib contrib-type="author" corresp="yes"><contrib-id contrib-id-type="orcid">https://orcid.org/0000-0002-6615-733X</contrib-id><name-alternatives><name name-style="eastern" xml:lang="ru"><surname>Лобанов</surname><given-names>А. А.</given-names></name><name name-style="western" xml:lang="en"><surname>Lobanov</surname><given-names>A. A.</given-names></name></name-alternatives><bio xml:lang="ru"><p>Москва</p></bio><bio xml:lang="en"><p>Moscow</p></bio><xref ref-type="aff" rid="aff-3"/></contrib><contrib contrib-type="author" corresp="yes"><contrib-id contrib-id-type="orcid">https://orcid.org/0000-0002-5616-5897</contrib-id><name-alternatives><name name-style="eastern" xml:lang="ru"><surname>Андронов</surname><given-names>С. В.</given-names></name><name name-style="western" xml:lang="en"><surname>Andronov</surname><given-names>S. V.</given-names></name></name-alternatives><bio xml:lang="ru"><p>Москва</p><p>Томск</p></bio><bio xml:lang="en"><p>Moscow</p><p>Tomsk</p></bio><xref ref-type="aff" rid="aff-4"/></contrib></contrib-group><aff-alternatives id="aff-1"><aff xml:lang="ru"><institution>Тюменский государственный университет</institution><country>Россия</country></aff><aff xml:lang="en"><institution>University of Tyumen</institution><country>Russian Federation</country></aff></aff-alternatives><aff-alternatives id="aff-2"><aff xml:lang="ru"><institution>Институт криосферы Земли ТюмНЦ СО РАН; НИУ Высшая школа экономики</institution><country>Россия</country></aff><aff xml:lang="en"><institution>Earth Cryosphere Institute, Tyumen Scientific Center SB RAS; HSE University</institution><country>Russian Federation</country></aff></aff-alternatives><aff-alternatives id="aff-3"><aff xml:lang="ru"><institution>Национальный медицинский исследовательский центр реабилитации и курортологии</institution><country>Россия</country></aff><aff xml:lang="en"><institution>National Medical Research Center for Rehabilitation and Balneology</institution><country>Russian Federation</country></aff></aff-alternatives><aff-alternatives id="aff-4"><aff xml:lang="ru"><institution>ФИЦ питания, биотехнологии и безопасности пищи; Томский государственный университет</institution><country>Россия</country></aff><aff xml:lang="en"><institution>Research Center for Nutrition, Biotechnology, and Food Safety; Tomsk State University</institution><country>Russian Federation</country></aff></aff-alternatives><pub-date pub-type="collection"><year>2026</year></pub-date><pub-date pub-type="epub"><day>09</day><month>07</month><year>2026</year></pub-date><volume>72</volume><issue>2</issue><fpage>236</fpage><lpage>251</lpage><permissions><copyright-statement>Copyright &amp;#x00A9; Соромотин А.В., Московченко Д.В., Федоров Р.Ю., Лобанов А.А., Андронов С.В., 2026</copyright-statement><copyright-year>2026</copyright-year><copyright-holder xml:lang="ru">Соромотин А.В., Московченко Д.В., Федоров Р.Ю., Лобанов А.А., Андронов С.В.</copyright-holder><copyright-holder xml:lang="en">Soromotin A.V., Moskovchenko D.V., Fedorov R.Y., Lobanov A.A., Andronov S.V.</copyright-holder><license xml:lang="ru" license-type="creative-commons-attribution" xlink:href="https://creativecommons.org/licenses/by/4.0/" xlink:type="simple"><license-p>Данная работа распространяется под лицензией Creative Commons Attribution 4.0.</license-p></license><license xml:lang="en" license-type="creative-commons-attribution" xlink:href="https://creativecommons.org/licenses/by/4.0/" xlink:type="simple"><license-p>This work is licensed under a Creative Commons Attribution 4.0 License.</license-p></license></permissions><self-uri xlink:href="https://www.aaresearch.science/jour/article/view/858">https://www.aaresearch.science/jour/article/view/858</self-uri><abstract><p>В исторической ретроспективе рассмотрены особенности социально-экологических взаимодействий, связанных с регулированием нагрузки на пастбищные угодья в оленеводстве на территории Ямало-Ненецкого автономного округа (ЯНАО). Установлено, что в период со второй половины XIX в. до начала существования советского государства традиционные формы урегулирования численности поголовья и маршрутов выпаса оленей способствовали сохранению высокого потенциала регенерации пастбищных угодий. В советский период постепенно стали возникать предпосылки к перевыпасу оленей, которые с 1960-х гг. проявились на территории Ямальского и Приуральского районов. Начиная с 1990- х гг. проблема перевыпаса оленей стала в той или иной мере характерной для всей территории ЯНАО. Характеризуя причины современной проблемы перевыпаса оленей, можно сделать вывод о том, что на нее большое влияние оказывают сокращение маршрутов выпаса и увеличение поголовья их стад на фоне сокращения угодий в процессе развития инфраструктуры нефтегазодобывающего комплекса. В этой ситуации необходима выработка новых подходов к регулированию оленеводства на основе разумного баланса между развитием его коммерческих функций и воздействием на арктические экосистемы.</p></abstract><trans-abstract xml:lang="en"><p>Against the backdrop of the reindeer herding crisis in many Arctic regions, the dynamics of its development in the north of Western Siberia are an exception. Over the past decade of the 20th century and the beginning of the 21st century, the territory of its largest reindeer herding center, the Yamalo-Nenets Autonomous Okrug of the Tyumen Region, has seen an increase in the number of reindeer, which currently accounts for about a third of the global total. Therefore this region can be considered a unique model territory which allows us to assess the nature of socio-natural interactions related to reindeer herding in their most intense form. In the historical retrospective, the paper examines the features of socio-natural interactions related to the regulation of the load on pasture lands in reindeer herding in the Yamalo-Nenets Autonomous Okrug. The study is based on the analysis of literature, archival sources, and the generalization of the authors' field research. It has been established that during the period from the second half of the 19th century to the beginning of the Soviet era, the traditional forms of regulating the number of reindeer and their grazing routes contributed to the preservation of a high potential for the regeneration of pasture lands. During the Soviet period, conditions gradually emerged for overgrazing of reindeer, and by the 1960s, it had become a problem in the Yamal and Priural districts. Since the 1990s, overgrazing of reindeer has become a serious issue. As to the causes of the problem, it has been established that the major factors are the reduction of grazing routes and an increase in the number of the herds against the background of the reduction of the land in the process of developing the infrastructure of the oil and gas production complex. Economic processes play a crucial role as factors that transform the Arctic ecosystems. The increasing irreversible negative consequences of overgrazing on the ecosystems could critically increase the vulnerability of reindeer herding. In this situation, it is necessary to develop new approaches to regulating reindeer herding based on achieving a reasonable balance between the development of its commercial functions and its impact on the Arctic ecosystems.</p></trans-abstract><kwd-group xml:lang="ru"><kwd>северное оленеводство</kwd><kwd>влияние оленеводства на экосистемы Субарктики</kwd><kwd>нагрузка на оленьи пастбища</kwd><kwd>проблема перевыпаса оленей</kwd><kwd>ЯНАО</kwd></kwd-group><kwd-group xml:lang="en"><kwd>northern reindeer herding</kwd><kwd>impact of reindeer herding on Subarctic ecosystems</kwd><kwd>load on reindeer pastures</kwd><kwd>problem of overgrazing</kwd><kwd>Yamalo-Nenets Autonomous Okrug</kwd></kwd-group><funding-group><funding-statement xml:lang="ru">Исследование выполнено за счет гранта Российского научного фонда № 24-16-00163, https://rscf.ru/project/24-16-00163/.</funding-statement><funding-statement xml:lang="en">The research has been supported by the Russian Science Foundation No. 24-16-00163, https://rscf.ru/en/project/24-16-00163/.</funding-statement></funding-group></article-meta></front><back><ref-list><title>References</title><ref id="cit1"><label>1</label><citation-alternatives><mixed-citation xml:lang="ru">Клоков К.Б. Прошлое и будущее оленеводства в этнокультурных ландшафтах коренных народов Арктики: анализ региональных различий. Арктика XXI век. Гуманитарные науки. 2023;1(31):54–62. https://doi.org/10.25587/SVFU.2023.47.95.006</mixed-citation><mixed-citation xml:lang="en">Klokov K.B. The past and future of reindeer herding in the ethnocultural landscapes of the indigenous peoples of the Arctic: an analysis of regional differences. The Arctic in the 21st Century. Humanities. 2023;1(31):54–62. (In Russ.). https://doi.org/10.25587/SVFU.2023.47.95.006</mixed-citation></citation-alternatives></ref><ref id="cit2"><label>2</label><citation-alternatives><mixed-citation xml:lang="ru">Кряжимский Ф.В., Маклаков К.В., Морозова Л.М., Эктова С.Н. Системный анализ биогеоценозов полуострова Ямал: имитационное моделирование воздействия крупностадного оленеводства на растительный покров. Экология. 2011;5:323–333.</mixed-citation><mixed-citation xml:lang="en">Kryazhimskij F.V., Maklakov K.V., Morozova L.M., Ektova S.N. System analysis of the biogeocenoses of the Yamal Peninsula: simulation modeling of the impact of large-scale reindeer herding on the vegetative cover. Ekologiya. 2011;5:323–333. (In Russ.).</mixed-citation></citation-alternatives></ref><ref id="cit3"><label>3</label><citation-alternatives><mixed-citation xml:lang="ru">Зуев С.М. Оленеводство в Ямало-Ненецком автономном округе: перспективы и проблемы. Научный вестник Ямало-Ненецкого автономного округа. 2015;3(88);103–107.</mixed-citation><mixed-citation xml:lang="en">Zuev S.M. Reindeer Herding in the Yamalo-Nenets Autonomous District: prospects and challenges. Scientific Bulletin of the Yamalo-Nenets Autonomous Okrug. 2015;3(88);103–107. (In Russ.).</mixed-citation></citation-alternatives></ref><ref id="cit4"><label>4</label><citation-alternatives><mixed-citation xml:lang="ru">Веселкин Д.В., Морозова Л.М., Горбунова А.М. Снижение значений NDVI в южных тундрах Ямала в 2001–2018 гг. коррелирует с численностью домашних северных оленей. Современные проблемы дистанционного зондирования Земли из космоса. 2021;18(2):143–155. https://doi.org/10.21046/2070-7401-2021-18-2-143-155</mixed-citation><mixed-citation xml:lang="en">Veselkin D.V., Morozova L.M., Gorbunova A.M. The decrease in NDVI values in the southern tundra of Yamal in 2001–2018 correlates with the number of domesticated reindeer. Sovremennye problemy distancionnogo zondirovaniya Zemli iz kosmosa. 2021;18:143–155. (In Russ.). https://doi.org/10.21046/2070-7401-2021-18-2-143-155</mixed-citation></citation-alternatives></ref><ref id="cit5"><label>5</label><citation-alternatives><mixed-citation xml:lang="ru">Yu Q., Epstein H.E., Frost G.V., Forbes B.C. Modeling dynamics of tundra plant communities on the Yamal Peninsula, Russia, in response to climate change and grazing pressure. Environ. Res. Lett. 2011;6:045505. https://doi.org/10.1088/1748-9326/6/4/045505</mixed-citation><mixed-citation xml:lang="en">Yu Q., Epstein H.E., Frost G.V., Forbes B.C. Modeling dynamics of tundra plant communities on the Yamal Peninsula, Russia, in response to climate change and grazing pressure. Environ. Res. Lett. 2011;6:045505. https://doi.org/10.1088/1748-9326/6/4/045505</mixed-citation></citation-alternatives></ref><ref id="cit6"><label>6</label><citation-alternatives><mixed-citation xml:lang="ru">Волковицкий А.И., Терехина А.Н. Современные проблемы ямальского оленеводства: дискуссии и перспективы. Этнография. 2020;2(8):152–169. https://doi.org/10.31250/2618-8600-2020-2(8)-152-169</mixed-citation><mixed-citation xml:lang="en">Volkovitskiy A., Terekhina A. The contemporary issues of Yamal reindeer herding: discussions and perspectives. Etnografia. 2020; 2(8):152-169. (In Russ.). https://doi.org/10.31250/2618-8600-2020-2(8)-152-169</mixed-citation></citation-alternatives></ref><ref id="cit7"><label>7</label><citation-alternatives><mixed-citation xml:lang="ru">Головнев А.В., Абрамов И.В. Олени и газ: стратегии развития Ямала. Вестник археологии, антропологии и этнографии. 2014;4(27):122–131.</mixed-citation><mixed-citation xml:lang="en">Golovnev A.V., Abramov I.V. Reindeer and gas: Yamal’s development strategies. Vestnik arheologii, antropologii i etnografii. 2014;4(27):122–131. (In Russ.).</mixed-citation></citation-alternatives></ref><ref id="cit8"><label>8</label><citation-alternatives><mixed-citation xml:lang="ru">Южаков А.А., Мухачев А.Д. Этническое оленеводство Западной Сибири: ненецкий тип. Новосибирск: Наука; 2001. 112 с.</mixed-citation><mixed-citation xml:lang="en">Южаков А.А., Мухачев А.Д. Этническое оленеводство Западной Сибири: ненецкий тип. Новосибирск: Наука; 2001. 112 с.</mixed-citation></citation-alternatives></ref><ref id="cit9"><label>9</label><citation-alternatives><mixed-citation xml:lang="ru">Крупник И.И. Арктическая этноэкология. М.: Наука; 1989. 272 с.</mixed-citation><mixed-citation xml:lang="en">Крупник И.И. Арктическая этноэкология. М.: Наука; 1989. 272 с.</mixed-citation></citation-alternatives></ref><ref id="cit10"><label>10</label><citation-alternatives><mixed-citation xml:lang="ru">Головнев А.В. Историческая типология хозяйства народов Севера Западной Сибири. Новосибирск: Изд-во Новосибирского государственного университета; 1993. 204 с.</mixed-citation><mixed-citation xml:lang="en">Головнев А.В. Историческая типология хозяйства народов Севера Западной Сибири. Новосибирск: Изд-во Новосибирского государственного университета; 1993. 204 с.</mixed-citation></citation-alternatives></ref><ref id="cit11"><label>11</label><citation-alternatives><mixed-citation xml:lang="ru">Народы Западной Сибири: Ханты. Манси. Селькупы. Ненцы. Энцы. Нганасаны. Кеты. М.: Наука; 2005. 805 с.</mixed-citation><mixed-citation xml:lang="en">Народы Западной Сибири: Ханты. Манси. Селькупы. Ненцы. Энцы. Нганасаны. Кеты. М.: Наука; 2005. 805 с.</mixed-citation></citation-alternatives></ref><ref id="cit12"><label>12</label><citation-alternatives><mixed-citation xml:lang="ru">Харючи Г.П. Традиции и инновации в культуре ненецкого этноса. Томск: Изд-во Томского ун-та; 2001. 228 с.</mixed-citation><mixed-citation xml:lang="en">Харючи Г.П. Традиции и инновации в культуре ненецкого этноса. Томск: Изд-во Томского ун-та; 2001. 228 с.</mixed-citation></citation-alternatives></ref><ref id="cit13"><label>13</label><citation-alternatives><mixed-citation xml:lang="ru">Форбс Б., Штаммлер Ф., Кумпула Т., Месштыб Н., Паюнен А., Каарлеярви В. Ямальские оленеводы, газодобыча и изменение окружающей среды: Адаптивный потенциал кочевого хозяйства и его ограничения. Экологическое планирование и управление. 2011;1:52–68.</mixed-citation><mixed-citation xml:lang="en">Forbs B., Shtammler F., Kumpula T., Messhtyb N., Payunen A., Kaarleyarvi V. Yamal reindeer herders, gas production, and environmental change: the adaptive potential of nomadic farming and its limitations. Ekologicheskoe planirovanie i upravlenie. 2011;1:52–68. (In Russ.).</mixed-citation></citation-alternatives></ref><ref id="cit14"><label>14</label><citation-alternatives><mixed-citation xml:lang="ru">Degteva A., Nellemann C. Nenets migration in the landscape: impacts of industrial development in Yamal peninsula, Russia. Pastoralism. 2013;3:15. https://doi.org/10.1186/2041-7136-3-15</mixed-citation><mixed-citation xml:lang="en">Degteva A., Nellemann C. Nenets migration in the landscape: impacts of industrial development in Yamal peninsula, Russia. Pastoralism. 2013;3:15. https://doi.org/10.1186/2041-7136-3-15</mixed-citation></citation-alternatives></ref><ref id="cit15"><label>15</label><citation-alternatives><mixed-citation xml:lang="ru">Клоков К.Б., Михайлов В.В. Выявление территорий климатического оптимума для традиционного оленеводства коренных народов Ямало-Ненецкого автономного округа. Известия Санкт-Петербургского государственного аграрного университета. 2015;40:105–108.</mixed-citation><mixed-citation xml:lang="en">Klokov K.B., Mihajlov V.V. Identification of climatic optimum areas for traditional reindeer herding by the indigenous peoples of the Yamalo-Nenets Autonomous District. Izvestiya Sankt-Peterburgskogo gosudarstvennogo agrarnogo universiteta. 2015;40:105–108. (In Russ.).</mixed-citation></citation-alternatives></ref><ref id="cit16"><label>16</label><citation-alternatives><mixed-citation xml:lang="ru">Клоков К.Б. Оленеводческие ландшафты России: ландшафтное районирование и траектории эволюции оленеводческого хозяйства в конце ХХ — начале ХХI столетия. Сибирские исторические исследования. 2023;3:96–112. https://doi.org/10.17223/2312461X/41/6</mixed-citation><mixed-citation xml:lang="en">Klokov K.B. Reindeer herding landscapes of Russia: landscape zoning and evolution trajectories of reindeer herding in the late 20th and early 21st centuries. Sibirskie istoricheskie issledovaniya. 2023;3:96–112. (In Russ.). https://doi.org/10.17223/2312461X/41/6</mixed-citation></citation-alternatives></ref><ref id="cit17"><label>17</label><citation-alternatives><mixed-citation xml:lang="ru">Истомин К.В., Хабек Й.О. Почвы криолитозоны и традиционное природопользование коренного населения северо-востока европейской части России и Западной Сибири: постановка исследовательской проблемы. Вестник археологии, антропологии и этнографии. 2019;1(44):108–119. https://doi.org/10.20874/2071-0437-2019-44-1-108-119</mixed-citation><mixed-citation xml:lang="en">Istomin K.V., Habek J.O. Soils of the cryolithozone and traditional nature management of the indigenous population of the Northeast of European Russia and Western Siberia: formulation of a research problem. Vestnik arheologii, antropologii i etnografii. 2019;1(44):108–119. (In Russ.). https://doi.org/10.20874/2071-0437-2019-44-1-108-119</mixed-citation></citation-alternatives></ref><ref id="cit18"><label>18</label><citation-alternatives><mixed-citation xml:lang="ru">Miles V. Esau I. Spatial heterogeneity of greening and browning between and within bioclimatic zones in northern West Siberia. Environ. Res. Lett. 2016;11:115002. https://doi.org/10.1088/1748-9326/11/11/115002</mixed-citation><mixed-citation xml:lang="en">Miles V. Esau I. Spatial heterogeneity of greening and browning between and within bioclimatic zones in northern West Siberia. Environ. Res. Lett. 2016;11:115002. https://doi.org/10.1088/1748-9326/11/11/115002</mixed-citation></citation-alternatives></ref><ref id="cit19"><label>19</label><citation-alternatives><mixed-citation xml:lang="ru">Советское оленеводство. Вып. 1: Кормовые ресурсы тундровой зоны СССР. Л.: Ин-т оленеводства; 1934. 211 с.</mixed-citation><mixed-citation xml:lang="en">Советское оленеводство. Вып. 1: Кормовые ресурсы тундровой зоны СССР. Л.: Ин-т оленеводства; 1934. 211 с.</mixed-citation></citation-alternatives></ref><ref id="cit20"><label>20</label><citation-alternatives><mixed-citation xml:lang="ru">Востряков П.Н., Броднев М.М. Оленеводство Ямала. Свердловск: Средне-Уральское книжное издательство; 1964. 98 с.</mixed-citation><mixed-citation xml:lang="en">Востряков П.Н., Броднев М.М. Оленеводство Ямала. Свердловск: Средне-Уральское книжное издательство; 1964. 98 с.</mixed-citation></citation-alternatives></ref><ref id="cit21"><label>21</label><citation-alternatives><mixed-citation xml:lang="ru">Лавренко Е.М., Корчагина А.А. (ред.). Полевая геоботаника. Т.4. Л.: Наука; 1972. 336 с.</mixed-citation><mixed-citation xml:lang="en">Лавренко Е.М., Корчагина А.А. (ред.). Полевая геоботаника. Т.4. Л.: Наука; 1972. 336 с.</mixed-citation></citation-alternatives></ref><ref id="cit22"><label>22</label><citation-alternatives><mixed-citation xml:lang="ru">Golovatin M.G., Morozova L.M., Ektova S.N., Paskhalny S.P. The change of tundra biota at Yamal peninsula (the North of the Western Siberia, Russia) in connection with anthropogenic and climatic shifts. In: Tundras: Wildlife and Climate trends. New York: Nova Science Publishers; 2010. P. 1–46.</mixed-citation><mixed-citation xml:lang="en">Golovatin M.G., Morozova L.M., Ektova S.N., Paskhalny S.P. The change of tundra biota at Yamal peninsula (the North of the Western Siberia, Russia) in connection with anthropogenic and climatic shifts. In: Tundras: Wildlife and Climate trends. New York: Nova Science Publishers; 2010. P. 1–46.</mixed-citation></citation-alternatives></ref><ref id="cit23"><label>23</label><citation-alternatives><mixed-citation xml:lang="ru">Богданов В.Д., Головатин М.Г., Морозова Л.М., Эктова С.Н. Социально-экологические условия промышленного освоения полуострова Ямал. Экономика региона. 2012;3(31):141–150. https://doi.org/10.17059/2012-3-13</mixed-citation><mixed-citation xml:lang="en">Bogdanov V.D., Golovatin M.G., Morozova L.M., Ektova S.N. Social and environmental conditions of industrial development in the Yamal Peninsula. Ekonomika regiona. 2012;3(31):141–150. (In Russ.). https://doi.org/10.17059/2012-3-13</mixed-citation></citation-alternatives></ref><ref id="cit24"><label>24</label><citation-alternatives><mixed-citation xml:lang="ru">Forbes B.C., Fauria M.M., Zetterberg P. Russian Arctic warming and ‘greening’ are closely tracked by tundra shrub willows. Global Change Biology. 2010:16;1542–1554. https://doi.org/10.1111/j.1365-2486.2009.02047.x</mixed-citation><mixed-citation xml:lang="en">Forbes B.C., Fauria M.M., Zetterberg P. Russian Arctic warming and ‘greening’ are closely tracked by tundra shrub willows. Global Change Biology. 2010:16;1542–1554. https://doi.org/10.1111/j.1365-2486.2009.02047.x</mixed-citation></citation-alternatives></ref><ref id="cit25"><label>25</label><citation-alternatives><mixed-citation xml:lang="ru">Frost G.V, Epstein, H.E., Walker, D.A. Regional and landscape-scale variability of Landsat-observed vegetation dynamics in northwest Siberian tundra. Environ. Res. Lett. 2014;9:025004. https://doi.org/10.1088/1748-9326/9/2/025004</mixed-citation><mixed-citation xml:lang="en">Frost G.V, Epstein, H.E., Walker, D.A. Regional and landscape-scale variability of Landsat-observed vegetation dynamics in northwest Siberian tundra. Environ. Res. Lett. 2014;9:025004. https://doi.org/10.1088/1748-9326/9/2/025004</mixed-citation></citation-alternatives></ref><ref id="cit26"><label>26</label><citation-alternatives><mixed-citation xml:lang="ru">Kitti H., Forbes B.C., Oksanen J. Long-and short-term effects of reindeer grazing on 314 tundra wetland vegetation. Polar Biology. 2009;32:253–261. https://doi.org/10.1007/s00300-008-0526-9</mixed-citation><mixed-citation xml:lang="en">Kitti H., Forbes B.C., Oksanen J. Long-and short-term effects of reindeer grazing on 314 tundra wetland vegetation. Polar Biology. 2009;32:253–261. https://doi.org/10.1007/s00300-008-0526-9</mixed-citation></citation-alternatives></ref><ref id="cit27"><label>27</label><citation-alternatives><mixed-citation xml:lang="ru">Письмаркина Е.В., Хитун О.В. Чужеродные виды растений в Ямало-Ненецком автономном округе (Россия): дополнения к флоре. Turczaninowia. 2019;22(1):26–34. https://doi. org/10.14258/turczaninowia.22.1.3</mixed-citation><mixed-citation xml:lang="en">Pis’markina E.V., Hitun O.V. Alien plant species in the Yamalo-Nenets Autonomous Okrug (Russia): additions to the flora. Turczaninowia. 2019;22(1):26–34. (In Russ.). https://doi. org/10.14258/turczaninowia.22.1.3</mixed-citation></citation-alternatives></ref><ref id="cit28"><label>28</label><citation-alternatives><mixed-citation xml:lang="ru">Горбунова А.М., Горбунов Л.С., Веселкин Д.В. Изменение запасов зеленых и лишайниковых кормов в сообществах южных тундр Ямала с 1930-х гг. по 2017–2019 гг. Экология. 2023;2:83–93. https://doi.org/10.31857/S0367059723020063</mixed-citation><mixed-citation xml:lang="en">Gorbunova A.M., Gorbunov L.S., Veselkin D.V. Changes in the availability of green and lichen-based forage in the southern tundra communities of Yamal from the 1930s to 2017–2019. Ekologiya. 2023;2:83–93. (In Russ.). https://doi.org/10.31857/S0367059723020063</mixed-citation></citation-alternatives></ref><ref id="cit29"><label>29</label><citation-alternatives><mixed-citation xml:lang="ru">Egelkraut D., Barthelemy H., Olofsson J. Reindeer trampling promotes vegetation changes in tundra heathlands: Results from a simulation experiment. J Veg Sci. 2020;31:476–486. https://doi.org/10.1111/jvs.12871</mixed-citation><mixed-citation xml:lang="en">Egelkraut D., Barthelemy H., Olofsson J. Reindeer trampling promotes vegetation changes in tundra heathlands: Results from a simulation experiment. J Veg Sci. 2020;31:476–486. https://doi.org/10.1111/jvs.12871</mixed-citation></citation-alternatives></ref><ref id="cit30"><label>30</label><citation-alternatives><mixed-citation xml:lang="ru">Морозова Л.М., Малыгина Н.В. Состояние растительного покрова тундровых пастбищ в районах многолетнего выпаса домашних и диких северных оленей. В мире научных открытий. 2013;73(43):49–79.</mixed-citation><mixed-citation xml:lang="en">Morozova L.M., Malygina N.V. The state of the vegetation cover of tundra pastures in areas of long-term grazing by domestic and wild reindeer. V mire nauchnyh otkrytij. 2013;43:49–79. (In Russ.).</mixed-citation></citation-alternatives></ref></ref-list><fn-group><fn fn-type="conflict"><p>The authors declare that there are no conflicts of interest present.</p></fn></fn-group></back></article>
